Table 1

Observed components of variance for ten types of relatives.
Observed component (y_i)	Method and types of relatives used to obtain component	Immune response	Body condition	Covariance

y₁	covariance between sire and offspring	0.003	0.106	0.019
y₂	covariance between nurse and offspring where the nurse is not the genetic dam	0.002	-0.122	-0.013
y₃	covariance between dam and offspring where the offspring was nursed by an unrelated dam	0.01	0.119	0.002
y₄	covariance between dam and offspring, where the dam is also the nurse.	0.006	0.022	0.022
y₅	covariance between full sibs raised by different nurses	0.010	0.328	0.025
y₆	covariance between unrelated sibs where the offspring were nursed by an unrelated dam	0.225	0.179	0.066
y₇	covariance between full sibs raised by different nurses	0.002	0.239	-0.002
y₈	covariance between unrelated sibs raised the same nurse	0.002	0.076	0.012
Y₉	covariance between unrelated sibs, each nursed by the genetic dam of the other	-0.215	0.149	-0.042
Y₁₀	variance among full sibs all with the same nurse	0.031	0.609	0.011

Definitions modified from Riska et al. [42]. Further details are in the Methods.
Gleeson et al. BMC Evolutionary Biology 2005 5:61 doi:10.1186/1471-2148-5-61