Schematic morphological drawing of the reproductive morphology of a land snail with one dart and a diverticulum. The love-dart (D) is produced and stored in the stylophore (S, often called dart sac) and shot by a forceful eversion of this organ. The mucus glands (MG) produce the mucus that is deposited on the dart before shooting. The penis (P) is intromitted to transfer the spermatophore. The sperm container is formed in the epiphallus (EP), while the spermatophore's tail is formed by the flagellum (FL). When a bursa tract diverticulum (BTD) is present, the spermatophore is received in this organ. Together with the bursa tract (BT) and bursa copulatrix (BC) these form the spermatophore-receiving organ (SRO, indicated in grey), which digest sperm and spermatophores. Sperm swim out via the tail of the spermatophore to enter the female tract and reach the sperm storage organ (SP, spermathecae) within the fertilization pouch (FP)-spermathecal complex. Other abbreviations: AG, albumen gland; G, genital pore; HD, hermaphroditic duct; OT, ovotestis; PRM, penis retractor muscle; SO, spermoviduct; V, vaginal duct; VD, vas deferens.
Koene and Schulenburg BMC Evolutionary Biology 2005 5:25 doi:10.1186/1471-2148-5-25