Maximum parsimony reconstruction of habitat type (lake versus river) using MacClade (the maximum likelihood reconstruction with Mesquite revealed analogous results) based on the maximum likelihood topology (see Fig. 3). The ancestor of the modern haplochromines (marked by a circle) is likely to have been a riverine species. This implies that also the Tanganyikan Tropheini originated from a riverine ancestor and re-colonized the lake where they presently form an abundant group in the rocky littoral zones.
Salzburger et al. BMC Evolutionary Biology 2005 5:17 doi:10.1186/1471-2148-5-17