Table 1

Individuals and sampling localities
morphospecies # locality geographical position EL
Carychium O.F. Müller, 1774 latitude longitude
C. clappi 1 USA, Tennessee, La Follette 36.332 −83.998 C1
Hubricht, 1959 2 USA, Tennessee, La Follette 36.332 −83.998 C1
3 USA, Tennessee, La Follette 36.332 −83.998 C1
4 USA, Tennessee, La Follette 36.332 −83.998 C1
C. exiguum 5 USA, New York, Naples, Ontario County, Grimes Glen 42.61591 −77.41355 C2
Say, 1822 6 USA, New York, Naples, Ontario County, Grimes Glen 42.61591 −77.41355 C2
7 USA, New York, Naples, Ontario County, Grimes Glen 42.61591 −77.41355 C2
C. exile 8 USA, Indiana, Lawrence County, Williams Cave Sinkhole 38.7555 −86.59105 C3
H. C. Lea, 1842 9 USA, Indiana, Lawrence County, Williams Cave Sinkhole 38.7555 −86.59105 C3
10 USA, Indiana, Lawrence County, Williams Cave Sinkhole 38.7555 −86.59105 C3
11 USA, New York, Portageville, Letchworth State Park 42.57909 −78.04945 C3
12 USA, New York, Portageville, Letchworth State Park 42.57909 −78.04945 C3
13 USA, New York, Watkins Glen 42.375828 −76.871115 C3
14 USA, New York, Watkins Glen 42.375828 −76.871115 C3
15 USA, New York, Watkins Glen 42.375828 −76.871115 C3
C. exile mexicanum 16 USA, Georgia, Adairsville, Bartow County, Barnsley Gardens 34.311233 −84.9866 C1
Pilsbry, 1891 17 USA, Georgia, Adairsville, Bartow County, Barnsley Gardens 34.311233 −84.9866 C1
18 USA, Alabama, Little River Mouth Park, Cherokee County 34.312233 −85.685733 C1
19 USA, Alabama, Little River Mouth Park, Cherokee County 34.312233 −85.685733 C1
20 USA, Illinois, Jackson County, Gorham 37.6865 −89.490667 C1
21 USA, Florida, Marianna 30.810556 −85.226667 C4
C. floridanum 22 USA, Florida, Wakulla Springs 30.23548 −84.303087 C5
Clapp, 1918 23 USA, Florida, Wakulla Springs 30.23548 −84.303087 C5
24 USA, Florida, Wakulla Springs 30.23548 −84.303087 C5
25 USA, Florida, Wakulla Springs 30.23548 −84.303087 C5
26 USA, Florida, Wakulla Springs 30.23548 −84.303087 C5
C. cf. schlickumi 27 Greece, Epirus, Ioáninna Prov., Métsovo 39.8213 21.1294 C6
Strauch, 1977 28 Greece, Epirus, Ioáninna Prov., Métsovo 39.8213 21.1294 C6
29 Greece, Epirus, Ioáninna Prov., Métsovo 39.8213 21.1294 C6
C. ibazoricum 30 Portugal, Azores, San Miguel, Sete Cidades 37.847033 −25.780217 C7
Bank & Gittenberger, 1985 31 Portugal, Azores, San Miguel, Furnas 37.770383 −25.306583 C7
32 Portugal, Azores, San Miguel, Furnas 37.770383 −25.306583 C7
33 Portugal, Estremadura, near Sao Pedro de Moel 39.774167 −9.016667 C7
34 Portugal, Estremadura, near Sao Pedro de Moel 39.774167 −9.016667 C7
C. lederi O. Boettger, 1880 35 Iran, Mazandaran, Nowshahr, Kheiroudkanar forest 36.605833 51.568333 C8
C. mexicanum 36 USA, Georgia, Flovilla, Butts County, Indian Springs State Park 33.242367 −83.92035 C9
Pilsbry, 1891 37 USA, Georgia, Flovilla, Butts County, Indian Springs State Park 33.242367 −83.92035 C9
38 USA, Georgia, Flovilla, Butts County, Indian Springs State Park 33.242367 −83.92035 C9
39 Belize, Maya Mountains, Bladen Nature Reserve 16.557167 −88.707833 C10
40 Belize, Maya Mountains, Bladen Nature Reserve 16.557167 −88.707833 C10
C. mexicanum costaricanum 41 Costa Rica, Puntarenas, Santa Elena 10.369833 −84.804167 C11
Von Martens, 1898 42 Costa Rica, Puntarenas, Monte Verde 10.301333 −84.790167 C11
43 Costa Rica, San José, San Gerardo de Dota 9.5496 −83.8032 C11
44 Costa Rica, San José, San Gerardo de Dota 9.5496 −83.8032 C11
45 Costa Rica, San José, San Gerardo de Dota 9.5496 −83.8032 C11
46 Panama, Chiriquí, Sendero el Retoño, Parque internacional la Amistad 8.8913 −82.628611 C12
47 Panama, Chiriquí, Sendero el Retoño, Parque internacional la Amistad 8.88815 −82.620667 C12
48 Panama, Chiriquí, Sendero el Retoño, Parque internacional la Amistad 8.88815 −82.620667 C12
49 Panama, Chiriquí, near Boquete 8.824767 −82.495833 C13
50 Panama, Chiriquí, near Boquete 8.824767 −82.495833 C13
51 Panama, Chiriquí, near Boquete 8.824767 −82.495833 C13
52 Panama, Chiriquí, near Boquete 8.824767 −82.495833 C13
C. minimum 53 Poland, Słubice 52.3467 14.5806 C14
O.F. Müller, 1774 54 Italy, Upper Adige, near Auer 46.3385 11.3503 C14
55 Spain, Asturias, Gijón 43.5204 −5.6164 C14
C. nannodes 56 USA, Tennessee, Overton County, Slit Cave 36.456833 −85.375067 C15
G.H. Clapp, 1905 57 USA, Tennessee, Unicoi County, near Unaka Springs 36.0986 −82.4466 C15
58 Canada, Ontario, Crawford Lake 43.4712 −79.9464 C15
C. nipponense 59 Japan, Honshu Is., Iwate, Ichinoseki, Geibikei 38.9845 141.255667 C16
Pilsbry & Hirase, 1904 60 Japan, Honshu Is., Fukushima, Urabandai 37.687783 140.1205 C16
C. noduliferum 61 Japan, Honshu Is., Iwate, Ichinoseki, Geibikei 38.9845 141.255667 C17
Reinhardt, 1877 62 Japan, Honshu Is., Fukushima, Urabandai 37.687783 140.1205 C17
63 Japan, Honshu Is., Kanagawa, Yamakita, Oomatazawa 35.423333 139.005 C18
64 Taiwan, Taichung County, Heping, Shei-Pa N.P., Huanshan 24.37295 121.310567 C19
C. occidentale 65 USA, Washington, Mason County 47.3038 −123.0918 C20
Pilsbry, 1891 66 USA, Washington, Mason County 47.3038 −123.0918 C20
67 USA, Washington, Mason County 47.3038 −123.0918 C20
68 USA, Washington, Mason County 47.3038 −123.0918 C20
69 USA, Washington, Mason County 47.3038 −123.0918 C20
C. cf. pessimum 70 Russia, Vladivostok, Promorsky Kray 43.193417 132.0511 C21
Pilsbry, 1902 71 Russia, Vladivostok, Promorsky Kray 43.193417 132.0511 C21
72 Russia, Vladivostok, Promorsky Kray 43.193417 132.0511 C21
73 Russia, Vladivostok, Promorsky Kray 43.193417 132.0511 C21
74 Russia, Vladivostok, Promorsky Kray 43.193417 132.0511 C21
75 Japan, Honshu Is., Iwate, Sarusawa 39.021167 141.292833 C22
C. stygium 76 USA, Kentucky, Hart County, Horse Cave (Hidden River Cave) 37.174167 −85.903833 C1
Call, 1897 77 USA, Kentucky, Hart County, Horse Cave (Hidden River Cave) 37.174167 −85.903833 C1
78 USA, Tennessee, Slit Cave (cave entrance) 36.456833 −85.375067 C1
79 USA, Tennessee, Slit Cave (cave entrance) 36.456833 −85.375067 C1
C. tridentatum 80 Switzerland, St. Gallen, St. Gallen 47.36 9.14 C23
(Risso, 1826) 81 Italy, Upper Adige, near Auer 46.3385 11.3503 C23
82 France, Britanny, near Lopreden 48.596667 −3.97 C23
Carychium sp. 1 83 Bulgaria, Mostovo Village, entrance Gargina Dupka Cave (Rhodopi Mnt.) 41.850667 24.926183 C23
84 Bulgaria, Mostovo Village, entrance Gargina Dupka Cave (Rhodopi Mnt.) 41.850667 24.926183 C23
85 Italy, Lombardy, Lago Maggiore, Laveno-Mombello 45.90718 8.6625 C23
86 Italy, Lombardy, Lago d'Iseo, Vigolo 45.7367 10.0025 C23
87 Romania, Bacâia 46.0249 23.1734 C23
88 Georgia, Kakheti Province, Lagodekhi Nature Reserve 41.88 46.31 C23
89 Georgia, Kakheti Province, Lagodekhi Nature Reserve 41.88 46.31 C23
Carychium sp. 2 90 China, Yunnan, Zhongdian County, Shangri-La 27.501017 100.033833 C24
91 China, Yunnan, Zhongdian County, Shangri-La 27.501017 100.033833 C24
92 China, Yunnan, Zhongdian County, Shangri-La 27.501017 100.033833 C24
93 China, Yunnan, Zhongdian County, Shangri-La 27.501017 100.033833 C24
94 China, Yunnan, Zhongdian County, Shangri-La 27.501017 100.033833 C24
Carychium sp. 3 95 China, North Sichuan, Songpan County, Huanglong 32.7373 103.824383 C25
96 China, North Sichuan, Songpan County, Huanglong 32.7373 103.824383 C25
97 China, North Sichuan, Songpan County, Huanglong 32.7373 103.824383 C25
Zospeum Bourguignat, 1856 cave name
Z. alpestre kupitzense 98 Slovenia, Kamnik-Savinja Alps, Solčava Ložekarjeva zijalka Z1
A. Stummer, 1984 99 Slovenia, Kamnik-Savinja Alps, Solčava Ložekarjeva zijalka Z1
Z. cf. biscaiense Gómez & Prieto, 1983 100 Spain, Garaimendi, Yurre Cueva de Otxas* Z2
Z. exiguum 101 Slovenia, Cerknica, Lož Križna jama* Z3
Kusčer, 1932 102 Slovenia, Cerknica, Lož Križna jama* Z3
103 Slovenia, Cerknica, Lož Križna jama* Z3
104 Slovenia, Cerknica, Lož Križna jama* Z3
105 Slovenia, Cerknica, Lož Križna jama* Z3
Z. frauenfeldi 106 Slovenia, Dobrepolje, Podpeč Podpeška jama* Z4
(Freyer, 1855) 107 Slovenia, Dobrepolje, Podpeč Podpeška jama* Z4
108 Slovenia, Dobrepolje, Podpeč Podpeška jama* Z4
109 Slovenia, Dobrepolje, Podpeč Podpeška jama* Z4
110 Slovenia, Dobrepolje, Podpeč Podpeška jama* Z4
Z. isselianum 111 Slovenia, Kamnik, Kamniška Bistrica Jama pod Farjevim plazom Z5
Pollonera, 1887 112 Slovenia, Kobarid*, Robič Turjeva jama Z6
113 Slovenia, Kamnik-Savinja Alps, Šmihel nad Mozirjem Konečka zijalka Z7
114 Slovenia, Kamnik-Savinja Alps, Šmihel nad Mozirjem Konečka zijalka Z7
115 Croatia, Kordun, Karlovac, Krnjak, Brebornica Mts. Jopićeva špilja Z8
116 Croatia, Kordun, Karlovac, Krnjak, Brebornica Mts. Jopićeva špilja Z8
117 Croatia, Kordun, Karlovac, Krnjak, Brebornica Mts. Jopićeva špilja Z8
118 Croatia, Kordun, Karlovac, Krnjak, Brebornica Mts. Jopićeva špilja Z8
Z. obesum 119 Slovenia, Gradiček, Krška vas Krška jama* Z9
(Frauenfeld, 1854) 120 Slovenia, Gradiček, Krška vas Krška jama* Z9
121 Slovenia, Gradiček, Krška vas Krška jama* Z9
Z. pretneri Bole, 1960 122 Croatia, Gračac, Kesići Donja Cerovačka špilja* Z10
Z. spelaeum schmidti 123 Slovenia, Veliki Otok, Postojna Betalov Spodmol Z11
(Frauenfeld, 1854) 124 Slovenia, Loka pri Mengšu Jama 1 pri Jabljah Z12
125 Slovenia, Loka pri Mengšu Jama 1 pri Jabljah Z12
126 Slovenia, Loka pri Mengšu Jama 1 pri Jabljah Z12
127 Slovenia, Loka pri Mengšu Jama 1 pri Jabljah Z12
128 Slovenia, Loka pri Mengšu Jama 1 pri Jabljah Z12
129 Slovenia, Krim Region, Gornji Ig Velika Pasica* Z11
130 Slovenia, Krim Region, Gornji Ig Velika Pasica* Z11
131 Italy, Trieste, near Gabrovizza San Primo Grotte d'Ercole Z11
132 Italy, Trieste, near Gabrovizza San Primo Grotte d'Ercole Z11
133 Italy, Trieste, near Basovizza Grotte Bac Z11
134 Italy, Trieste, near Basovizza Grotte Bac Z11
135 Italy, Trieste, near Samatorza Grotte Azzurra Z11
Z. suarezi 136 Spain, Cantabria, Novales, La Busta Cueva del Linar Z13
Gittenberger, 1980 137 Spain, Cantabria, Novales, La Busta Cueva del Linar Z13
138 Spain, Cantabria, Novales, La Busta Cueva del Linar Z13
139 Spain, Cantabria, Novales, La Busta Cueva del Linar Z13
140 Spain, Asturias, Inguanzo Cueva del Bosque/Cueva Inguanzo* Z14
141 Spain, Asturias, Inguanzo Cueva del Bosque/Cueva Inguanzo* Z14
142 Spain, Bizkaia, San Pedro de Galdames, Barranco de Aranaga Cueva de los Cuervos Z15
143 Spain, Bizkaia, San Pedro de Galdames, Barranco de Aranaga Cueva de los Cuervos Z15
144 Spain, Bizkaia, San Pedro de Galdames, Barranco de Aranaga Cueva de los Cuervos Z15
145 Spain, Castilla y León, Monte Santiago Cueva de Las Paúles Z16
Z. subobesum 146 Croatia, Ogulin, Tounj Tounjčica* Z8
Bole, 1974 147 Croatia, Ogulin, Tounj Tounjčica* Z8
148 Croatia, Ogulin, Tounj Tounjčica* Z8
Zospeum sp. 1 149 Spain, Bizkaia, Guipúzcoa Mts., Valle de Araotz Cueva de Ermita de Sandaili Z17
150 Spain, Bizkaia, Guipúzcoa Mts., Valle de Araotz Cueva de Ermita de Sandaili Z17
151 Spain, Bizkaia, Guipúzcoa Mts., Valle de Araotz Cueva de Ermita de Sandaili Z17
152 Spain, Garaimendi, Yurre Cueva de Otxas Z17
153 Spain, Garaimendi, Yurre Cueva de Otxas Z17
154 Spain, Garaimendi, Yurre Cueva de Otxas Z17
155 Spain, Garaimendi, Yurre Cueva de Otxas Z17
156 Spain, Garaimendi, Yurre Cueva de Otxas Z17
157 Spain, Bizkaia, Manaria, Urkuleta Valley Cueva Silibranka II Z17
158 Spain, Bizkaia, Manaria, Urkuleta Valley Cueva Silibranka II Z17
159 Spain, Bizkaia, Manaria, Urkuleta Valley Cueva Silibranka II Z17
160 Spain, Bizkaia, Dima, Indusi Cueva del Cráneo Z17
Zospeum sp. 2 161 Spain, Castilla y León, Monte Santiago Cueva de Las Paúles Z18
162 Spain, Castilla y León, Monte Santiago Cueva de Las Paúles Z18
163 Spain, Castilla y León, Monte Santiago Cueva de Las Paúles Z18
Outgroup taxa
Pythiidae, Laemodonta cubensis 164 Bermuda, Hamilton Parish, Walsingham Pond, outside Cliff Cave 32.34773 −64.70965
Melampodidae, Microtralia occidentalis 165 Bermuda, St. George Parish, St. George Island, Lover's Lake 32.36750 −64.70990
Veronicellidae, Veronicella cubensis 166 Bermuda, Devonshire Parish, Winfried Gibbons Nature Reserve, South Road 32.30230 −64.74450

Data on the morphological identification, taxonomic first description, specimen number (#), sampling locality and evolutionary lineage (EL) are provided. Type locality populations or localities referred to in the phenotype descriptions of Zospeum morphospecies are marked by an asterisk.

Weigand et al.

Weigand et al. BMC Evolutionary Biology 2013 13:18   doi:10.1186/1471-2148-13-18

Open Data