Table 1 

Information metrics of environmental complexity. 

Emergence of the organism with fitness w 

w > 0.75 
w >0.85 

Average speed, epochs 
Success Rate 
Average speed, epochs 
Success Rate 



Unevolved → AND 
39 
64/64 
236 
64/64 
Unevolved → OR 
1,179 
57/64 
> 4,000 
25/64 
Unevolved → A 
1,667 
60/64 
2,425 
59/64 
Unevolved → XOR 
5,241 
26/64 
9,583 
21/64 


AND → XOR 
4,891 
7/16 
> 6,000 
7/16 
XOR → AND 
250 
14/16 
1230 
6/16 


AND → OR 
> 4,000 
9/16 
 
2/16 
OR → AND 
17 
16/16 
199 
14/16 


A → AND 
125 
15/16 
696 
10/16 
AND → A 
1477 
14/16 
2362 
14/16 


A → OR 
220 
11/16 
2200 
10/16 
OR → A 
56 
16/16 
56 
16/16 


OR → XOR 
210 
15/16 
2093 
10/16 
XOR → OR 
42 
16/16 
119 
16/16 


A → XOR 
240 
14/16 
887 
11/16 
{A &B} → XOR + HGT 
138 
16/16 
423 
12/16 
XOR → A 
10 
16/16 
17 
16/16 


A populations is considered to be evolved once its fitness is above w = 0.75; additional phenotype refinement is required to pass the w = 0.85 threshold. Success rate is the ratio of simulations with fitness above threshold after 4,000 epochs over the total number of simulations. 

Mozhayskiy and Tagkopoulos BMC Bioinformatics 2012 13(Suppl 10):S10 doi:10.1186/1471210513S10S10 