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The multiple sex chromosomes of platypus and echidna are not completely identical and several share homology with the avian Z.
Rens W, O'Brien PC, Grützner F, Clarke O, Graphodatskaya D, Tsend-Ayush E, Trifonov VA, Skelton H, Wallis MC, Johnston S, Veyrunes F, Graves JA, Ferguson-Smith MA
Genome Biol 2007, 8:R243
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The status of dosage compensation in the multiple X chromosomes of the platypus.
Deakin JE, Hore TA, Koina E, Marshall Graves JA
PLoS Genet 2008, 4:e1000140
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