Spawning salmon disrupt trophic coupling between wolves and ungulate prey in coastal British Columbia

Chris T Darimont^1,2,4 , Paul C Paquet^2,3 and Thomas E Reimchen¹

¹  Department of Biology, Box 3020, Stn CSC, University of Victoria, Victoria, British Columbia, V8W 3N5, Canada,

²  Raincoast Conservation Foundation, Box 77, Denny Island, British Columbia, V0T 1B0, Canada,

³  Faculty of Environmental Design, Professional Faculties Building, Room 2182 University of Calgary, 2500 University Drive NW, Calgary, Alberta, T2N 1N4, Canada

⁴  Department of Environmental Studies, 405 ISB, University of California, 1156 High Street, Santa Cruz, California, 95060, USA