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Atsuki Hiyama, Chiyo Nohara, Wataru Taira, Seira Kinjo, Masaki Iwata and Joji M Otaki*
* Corresponding author: Joji M Otaki otaki@sci.u-ryukyu.ac.jp
BMC Evolutionary Biology 2013, 13:168 doi:10.1186/1471-2148-13-168
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