Figure 4.

Semi-logarithmic plot of Lineages Through Time for the Harpalinae clade only (A) and the Brachininae clade only (B). The dark black lines are the empirical chronograms from the BEAST analysis of all data combined. The lighter colored lines are simulations with incomplete taxon sampling generated under a pure birth process. Upturns and down turns in the empirical LTT plots reflect changes in rates of diversification. The arrows in the Harpaline LTT (A) show changes in diversification rates are estimated to have occurred according to the best fitting model of lineage diversification.